Among terrestrial vertebrates, the start of the Eocene is marked by the appearance of two new groups of animals: the perissodactyls, or odd-toed ungulates, and the artiodactyls, or even-toed ungulates. The perissodactyls include the horses, rhinoceroses, and tapirs; among the artiodactyls are the deer, cattle, and sheep. An early horse ancestor, the dawn horse, known in North America as Eohippus, is among the fossil perissodactyls found in the lower Eocene rocks of both North America and Europe. Artiodactyls, rare during the early Eocene, became abundant later in the epoch.
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